| Introduction
The studied
material originates from a formation situated in the vicinity of
the village of Satovcha (Sofia district) in the western extention
of the Rhodopes (Fig. 1). To be more concrete, it originates from
the Sivik Formation of the seams of Satovcha Grabben developed to
the south-east of the village. In the opinion of V a t s e v and
Piroumova (1983), the rocks of this formation date back to Middle-Upper
Miocene. In some publications devoted to different taxa from Satovcha
palaeoflora Palamarev, Bozukov (1992), Bozukov, Palamarev (1992),
B o z u-k o v (1995) suggest that to all probability these rocks
date from Middle Miocene (Badenian). This view is supported in the
present study, too.
The discovery of Davallia haidingeri E11.
is especially important both for purely paleofloristic studies and
for the reconstruction of palaeoecological conditions of the Western
Rhodopes during Middle Miocene. The contemporary representatives
of genus Davallia S m i t h are epiphytes spread in paleotropics
(T r y o n, L u g a r-d o n, 1990). This biological type has specific
requirements to the environment, such as relatively high temperature
and air humidity. Fossil epiphytes may serve as impor-tant indicator
species for the reconstruction of palaeoclimate. In view of this,
D. haidingeri E11., together with the other species from
the exceptionally rich Satovcha palaeoflora, will contribute to
more accurate reconstruction of palaeocological con-ditions.
The significance of this find lies in the expansion
of hitherto known stratigraphic and geographic areas.
The discovery of spores in the sori of a fossil
sample is of special importance. They are identical to those from
fossil species Polypodiisporites alienus (R. P o t. 1931)Nagy
1973. This fact sheds more light on the botanical affinity of the
fossil taxon.
Material and Methods
We have examined a leaf remain from a pinnate
leaf with sori No CAT-2419 from the Paleobotanical Collection of
the Institute of Botany, BAS. For the purpose ichnophytological
methods was applied (Z h i 1 i n, 1969, D i 1 c h e r, 1974).
For the preparation of palynological slides the
spores were isolated from the sori of the leaf remain and treated
with 10% HC1, 5% KOH and then acetolysis after E r d t m a n (1952).
Observations and microphotographs were made on glycerine-jelly microscope
preparation with Docuval photomicroscope at magnification of 500x
to 1500x. Morphological investigations and measurement of fossil
spores were performed on 30 spores arbitrarily chosen from 3 microscope
preparations. The processed material is preserved at the Palynological
Collection of the Institute of Botany, BAS.
The taxonomic revision byNoteboom (1994) for genus
Davallia was used for the purpose of the present study.
Results and Discussion
Family DA VALLIACEAE Frank
Genius Davallia Smith
Davallia haidingeriEttingshausen
PI. I, Fig. 1-5; PI. II, Fig. 1-7.
1858. Davallia haidingeri E 11 i n g s
h a u s e n, p. 50, PI. II, Fig. 5.
Description of the leaf remain: leaf impaired
pinnate, deltoid, with alternate or opposite partites of the 2nd
and 3rd order; partites of the 2nd order also impaired pinnate,
with 9-11 elongated partites of the 3rd order which in their turn
are in dented or lobated into 5-7 partites of the 5th-7th order.
The sori are situated close to the leaf margin in partites of the
3rd or the 4th order. Dimensions: leaf length - probably about 60.0
cm; length of the preserved partites of the 2nd order - 2.5-5.0
cm; width - 1.0-1.5 cm; length of the preserved partites of the
3rd order - 0.5-1.3 cm; width - 0.2-0.5 cm.
Comparison. No devations from the holotype were
observed in the examined sample (PI. I, Fig. I).
Ettingshausen (1858) compares this fossil species
with the recent species Davallia canariensis (L.) S m. (PI. IV,
Fig. 1) spread in Macaronesia, Maroco, the Canary Islands and the
Iberian Peninsula from 100 to 1000 m a.s.l.
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1.
Sketch-map of the area investigated
1.
Davallia haidingeri E11., - Sotzka, Slovenia, Holotypus (xl);
2.
Davallia haidingeriE11. - Satovcha Graben, No CAT-2419 (xl);
3.
Davallia haidingeriE 11. - Satovcha Graben, No CAT-2419. fragment
(x2);
4.
Davallia haidingeriE11. - Satovcha Graben, No CAT-2419, fragment
(x3);
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